Table 4. Emigration rates survival and dispersal-ratio

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Table VII-4. Estimates of the emigration rate, survival during dispersal and the dispersal-ratio
as derived by an iterative method best fitting the field data.

R = number of beetles released;

E_m = number of beetles emigrated;

I_e = number of emigrants found as immigrant elsewhere;

I = number of immigrants from elsewhere;

E = emigration rate/week;

Q_e = survival chance during dispersal;

Db = dispersal-ratio; v.c. = variation coefficient (standard deviation/mean)

BORDER POOLS CENTRE POOLS

POOL: Wn Bb Me Kr Br Wz Sc Kw Bn

period

R

I 46 70 68 - 49 84 41 68 -

II 70 58 60 - 25 70 70 - -

III 90 40 100 85 85 75 - - 87

IV 100 - 80 - 90 120 - - -

E_m

I 14 39 45 - 37 57 31 54 -

II 49 14 33 - 15 52 46 - -

III 45 15 28 43 19 58 - - 66

IV 32 - 21 - 44 42 - - -

I_e

I 12 23 26 - 33 40 16 41 -

II 38 5 7 - 10 45 37 - -

III 42 13 28 29 10 56 - - 62

IV 30 - 19 - 29 16 - - -

I

I 30 2 40 3 43 17 38 6 12

II 14 0 7 2 26 49 26 2 16

III 45 20 16 49 77 9 0 0 17

IV 6 1 16 2 11 31 24 0 3

BORDER POOLS CENTRE POOLS

POOL: Wn Bb Me Kr MEAN v.c. Br Wz Sc Kw Bn MEAN v.c. MEAN v.c.

E

I 0.13 0.18 0.58 - 0.30 0.83 0.50 0.27 0.42 0.45 - 0.41 0.24 0.36 0.47

II 0.32 0.07 0.19 - 0.19 0.65 0.30 0.38 0.35 - - 0.34 0.12 0.27 0.44

III 0.18 0.17 0.07 0.18 0.15 0.36 0.08 0.65 - - 0.50 0.41 0.72 0.26 0.85

IV 0.12 - 0.10 - 0.11 0.13 0.36 0.23 - - - 0.30 0.31 0.20 0.59

Q_e

I 0.86 0.58 0.58 - 0.67 0.24 0.89 0.70 0.52 0.77 - 0.72 0.21 0.70 0.21

II 0.78 0.35 0.21 - 0.45 0.67 0.68 0.86 0.81 - - 0.78 0.12 0.62 0.44

III 0.93 0.89 1.00 0.67 0.87 0.16 0.52 0.97 - - 0.93 0.81 0.31 0.84 0.21

IV 0.95 - 0.89 - 0.92 0.05 0.65 0.38 - - - 0.52 0.37 0.72 0.36

Db

I 2.16 0.05 0.89 - 1.03 1.03 1.15 0.30 1.23 0.11 - 0.70 0.82 0.97 0.76

II 0.29 0.00 0.21 - 0.17 0.90 1.76 0.94 0.57 - - 1.09 1.09 0.63 1.02

III 1.00 1.37 0.58 1.14 1.02 0.32 4.02 0.16 - - 0.26 1.48 1.49 1.22 1.08

IV 0.19 - 0.75 - 0.47 0.84 0.25 0.73 - - - 0.49 0.69 0.48 0.63

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Table VII-4.	Estimates of the emigration rate, survival during dispersal and the dispersal-ratio as derived by an iterative method best fitting the field data.
R =	number of beetles released;
E_m =	number of beetles emigrated;
I_e =	number of emigrants found as immigrant elsewhere;
I =	number of immigrants from elsewhere;
E =	emigration rate/week;
Q_e =	survival chance during dispersal;
Db =	dispersal-ratio; v.c. = variation coefficient (standard deviation/mean)

	BORDER POOLS						CENTRE POOLS
POOL:	Wn	Bb	Me	Kr			Br	Wz	Sc	Kw	Bn
period
R
I	46	70	68	-			49	84	41	68	-
II	70	58	60	-			25	70	70	-	-
III	90	40	100	85			85	75	-	-	87
IV	100	-	80	-			90	120	-	-	-
E_m
I	14	39	45	-			37	57	31	54	-
II	49	14	33	-			15	52	46	-	-
III	45	15	28	43			19	58	-	-	66
IV	32	-	21	-			44	42	-	-	-
I_e
I	12	23	26	-			33	40	16	41	-
II	38	5	7	-			10	45	37	-	-
III	42	13	28	29			10	56	-	-	62
IV	30	-	19	-			29	16	-	-	-
I
I	30	2	40	3			43	17	38	6	12
II	14	0	7	2			26	49	26	2	16
III	45	20	16	49			77	9	0	0	17
IV	6	1	16	2			11	31	24	0	3
	BORDER POOLS						CENTRE POOLS
POOL:	Wn	Bb	Me	Kr	MEAN	v.c.	Br	Wz	Sc	Kw	Bn	MEAN	v.c.	MEAN	v.c.
E
I	0.13	0.18	0.58	-	0.30	0.83	0.50	0.27	0.42	0.45	-	0.41	0.24	0.36	0.47
II	0.32	0.07	0.19	-	0.19	0.65	0.30	0.38	0.35	-	-	0.34	0.12	0.27	0.44
III	0.18	0.17	0.07	0.18	0.15	0.36	0.08	0.65	-	-	0.50	0.41	0.72	0.26	0.85
IV	0.12	-	0.10	-	0.11	0.13	0.36	0.23	-	-	-	0.30	0.31	0.20	0.59
Q_e
I	0.86	0.58	0.58	-	0.67	0.24	0.89	0.70	0.52	0.77	-	0.72	0.21	0.70	0.21
II	0.78	0.35	0.21	-	0.45	0.67	0.68	0.86	0.81	-	-	0.78	0.12	0.62	0.44
III	0.93	0.89	1.00	0.67	0.87	0.16	0.52	0.97	-	-	0.93	0.81	0.31	0.84	0.21
IV	0.95	-	0.89	-	0.92	0.05	0.65	0.38	-	-	-	0.52	0.37	0.72	0.36
Db
I	2.16	0.05	0.89	-	1.03	1.03	1.15	0.30	1.23	0.11	-	0.70	0.82	0.97	0.76
II	0.29	0.00	0.21	-	0.17	0.90	1.76	0.94	0.57	-	-	1.09	1.09	0.63	1.02
III	1.00	1.37	0.58	1.14	1.02	0.32	4.02	0.16	-	-	0.26	1.48	1.49	1.22	1.08
IV	0.19	-	0.75	-	0.47	0.84	0.25	0.73	-	-	-	0.49	0.69	0.48	0.63